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Predators. . .
                                                    Around the world, predators are practicing
                                                    survival of the fittest. This surviving has been in
                                                    action for more than a million years for most
                                                    species. However, extinction rates post human-
                                                    introduction have multiplied by the hundreds,
                                                    possibly thousands.
                                                    This question, then, is posed: Why are so many
                                                    species now disappearing at an accelerated
                                                    pace?

The sleek, sheen coat of the              With the incredible shrinking      ecosystem. To understand why
cougar might for a moment             populations of predator species,       each element is so crucial to
distract you from the glistening      scientists find them difficult to      biodiversity, let’s recall the food
canines capable of ripping flesh      study. These predators are             chain.
from bone. Sunning at the zoo,        reduced to fractions of their              Every living organism in an
these ‘big cats’ hardly appear to     former habitats and are                ecosystem supports other living
retain their natural predatory        instinctively secretive; it is not     organisms. For example, in Figure
ways. Here, safely locked away        surprising that their sightings are    1 the vegetation supports the
behind thick glass walls, they        rare. Without ample research           rabbit. The
stimulate our curiosity, our          subjects, scientists are quick to      rabbit in
interest, even perhaps our desire.    dismiss studies on predators.          turn
One can be captivated by the          However, in studying their             supports
power of the majestic, fluid          absence, we can still draw very        the wolf.
movements of these magnificent        important conclusions about these      The wolf,
creatures.                            species.                               after dying,
    Still, this awe and amazement                                            is turned
has done little to aid the cats in                                           into
                                     Top predators are essential to
surviving the onset of humans.                                               necessary
                                     the integrity of ecological
The cougar population, much like                                             nutrients in
that of the gray wolves, has                                                 the soil that
                                     communit ies… it is imperative
steadily declined with the                                                   will support
                                     to retain or restore top
increase of human populations.                                               the growth
Farmers, ranchers, and residents                                             of more
                                     predators to as many parts of
of these areas seem to have no                                               vegetation.
                                     North America as pract ical…
complaints with the disappearing                                             This is a
predators. That is until the prey                                            minimal         Figure 1
                                                          Terborgh, et al.
these predators had been keeping                                             representation; food chains are in
in check trample their gardens,                                              actuality much more complex and
eat their crops, and disease their        One such conclusion is that the    usually compound into a food web
herds.                                loss of top predators world-wide       (as each organism relies on more
    The prevalent eradication of      has upset the biodiversity of many     than one source for its survival).
top predators from ecosystems         ecosystems. Biodiversity is the        However, as you can see from this
has agitated the regulatory           balance between all organisms          simple depiction, removing one
balance these predators once          occupying an ecological                any single element will interrupt
provided. Our removal of              community. Varied vegetation,          the life-supporting role and the
predatory species has eliminated      herbivore grazing, and predation       balance in which each element
their role of regulating prey         all play important roles in            regulates the other elements. This
species’ populations.                 maintaining biodiversity in each       is called a trophic cascade.
We can find trophic cascades       cottonwoods along the                 herbivores onto predator-free
in both top-down and bottom-up         streambanks and heavy bank            islands by sailors centuries ago to
processes at work in ecosystems.       erosion…” The same effect has         ensure a food supply upon
“Top-down” stems from the levels       been noted in Yellowstone National successive visits. Without any
of the tropic chain (i.e., the food    Park (as demonstrated in the video natural predators to control the
chain) being manipulated from the      linked below) with the removal of     number of herbivores, the plant-
top of the chain down. To expand       wolves and the dramatic increase      eaters destroyed the vegetation of
on this, return to our previous        in the number of elks.                the islands. On islands like
example. Removing the wolf from            The amazing factor not yet        Ascension, Juan Fernandez, the
the simple food chain would allow      considered is how these trophic       California Channel Islands, and St.
the rabbit to overpopulate (as         cascades work in detriment to the     George Island, the herbivores’
rabbits are known to do). As a         species whose numbers explode.        destruction of the existing plant-
result, the vegetation the rabbit      They quickly diminish their own       life lead to their own downfall, as
population survives on would           resources, resources that also        they now had no resources on
suffer, unable to keep up with the     support other species. To explain,    which to survive.
increase of its superior trophic       think of the food supply that both
                                       white-tailed deer and rabbits rely Mammals that would have been
level’s increase. The opposite of
                                                                          part of the prey pool of missing
this effect is true in bottom-up       on. With the elimination of the
processes. If we were to increase      white-tailed deer’s main
                                                                          carnivores, such as wolves and
the number of plants available, this   predator, the deer will begin to
                                                                          cougars, have become notoriously
too could cause the rabbit             consume more of the food
                                       supply. Rabbits, in turn, are then abundant to the point that some of
population, and thus the wolf
population, to amplify.                left with less food and their
                                                                          them are now nuisances: by being
    Trophic cascades are being         population decreases. This
                                       cause-and-effect action is known road hazards, by browsing
recognized in national parks
across America; one such example       as the “Paine effect.” Through     ornamental shrubbery, by raiding
has been documented in The             the Paine effect, we can
                                                                          trashcans, by preying on birds and
Journal of Biological Conservation,    determine that removing the top
                                       predator of an ecosystem results their nests, by destroying vegetable
remarking on the loss of cougars in
Zion National Park. Professor          in loss of diversity of the prey   gardens, and by flooding people’s
Robert Beschta of Oregon State         community; again, we can
                                                                          yards.
University states, “When park          conclude that top predators are
                                                                                                     Terborgh, et al.
development caused cougars to          absolutely crucial to maintaining
begin leaving Zion Canyon in the       biodiversity (which is essential
1930s, it allowed much higher          for all species’ survival).                In the modern era, humans are
levels of deer browsing. That set in       Reviewing multiple ecosystems responsible for the missing top
motion a long cascade of changes       around the globe supports this        predators. In their absence, we
that resulted in the loss of most      conclusion. One such example was are noticing an explosion of their



More on Trophic Cascades
    The standard definition of
trophic is “of or relating to
nutrition.” Each level of the food
chain is a trophic level. A trophic
cascade is where different levels
of the food chain or web are
affected by the loss or addition of
another element. To see exactly
how trophic cascades work, visit:
http://youtube.com/watch?
v=O8BV0VCn1JU for a short
video.
former prey. Prey species’         caribou, elk, deer, and moose
                                                   populations expanding have         populations have declined.
                                                   caused a noticeable amount         While these examples have not
                                                   of problems for humans.            been controlled experiments
                                                   Deer-related car accidents         and not always carefully
                                                   number into the thousands          monitored, their results are
                                                   annually; moose, too, stray        consistent. Human interference
                                                   into highways at their active      in nature’s design, specifically
                                                   times, which happens after         the intricate roles of the trophic
                                                   dark when drivers’ visibility is   ladder, typically produces
                                                   already reduced. Raccoons          negative reactions throughout
                                                   and opossums are just as           the ecosystem for all species
                                                   responsible for overturned         involved.
                                                   trashcans as stray dogs;
                                                                                      The conclusion that top
                                                   beavers, misplaced from their
                                                   natural territories and without    predators play a major
                                                   predators to keep their
                                                                                      regulatory role seems
                                                   numbers in check, build dams
                                                                                      inescapable.
                                                   in the most inconvenient
                                                   places flooding residential                           Terborgh, et al.
                                                   yards. As frustrating as each
                                                   of these situations may be,            As more and more
                                                   people have no where to            environmental issues appear
                                                   place the blame outside of         on the screens of our
                                                   our own species. Our               televisions, the front pages of
                                                   reduction of natural habitats      our newspapers, and in the
                                                   and elimination of natural         headlines of our favorite online
                                                   predators has led to the           news, we cannot escape the
                                                   complications we are now           importance of conserving
                                                   experiencing, all a product of     nature. A top concern for
                                                   the loss of biodiversity.          conservationists is maintaining
                                                       We have also been known        biodiversity; one guaranteed
                                                   to upset biodiversity by           way to destabilize an
                                                   introducing predators into         ecosystem is by removing the
                                                   areas previously devoid of         top predators from it. Their
                                                   predators. When the brown          removal causes a chain
                                                   tree snake was introduced          reaction; first, we see an
                                                   and took up residence in           overabundance of targeted
                                                   Guam, it flourished.               prey. This consequently affects
                                                   Consequently, the native bird      other species who consume the
                                                   population drastically             same food as the now
                                                   decreased. With the                overabundant species. It also
                                                   reintroduction of the gray         disrupts and destroys
                                                   wolf in the northern Midwest       vegetation and habitats in the
                                                   area of the United States,         ecosystem. There is hope...

More on Ecosystems
An ecosystem is an interconnected community of
living organisms and the physical environment in
which they interact to form a life-support system.
You don’t have to travel to Yellowstone or a state
park to witness an ecosystem at work; you can see
one in your own backyard! Discovery Channel’s
Animal Planet hosts a show, Backyard Habitat,
which assists families with developing ecosystems at
their homes. Visit: http://animal.discovery.com/
fansites/backyard/backyard.html for example
habitats, tips on renovating your own yard, or to see
how to have your yard certified through the National
Wildlife Federation.
For more information:
    Despite the gray wolf’s                The answer to the opening
protected status as an                 question remains without a
Endangered Species and the             single, conclusive answer.
                                                                               Keystone Conservation
                                                                           •
proven need for the cougar in          Many factors contribute to the
                                                                                   http://www.keystoneconservation.us
                                                                               •
maintaining balance within             loss of species. However,
ecosystems, both are legally           some factors we can control.
hunted within several states           We must take direct action to           Center for Biological Diversity
                                                                           •
(Cougars are currently not on          restoring and maintaining top
                                                                                   http://www.biologicaldiversity.org
                                                                               •
the Threatened or                      predators because their
Endangered Species lists;              irreplaceable and critical
                                                                               Mountain Lion Foundation
however, their sightings are           roles inside ecosystems             •
so rare there is no national or        affects practically every other             http://www.mountainlion.org
                                                                               •
state-specific count of the            organism dependant upon
                                       that ecosystem. We are part
remaining cats in the wild).
                                                                               Wildlife Rescue & Rehabilitation, Inc.
                                                                           •
                                       of their ecosystems.
At this time, though, there are
                                                                                   http://www.wildlife-rescue.org
groups working to limit and                                                    •
hopefully, eventually remove
these species from open-                   In addition to visiting your
                                                                               Wolf Conservation Center
                                                                           •
game lists. They propose that          state’s official website for
                                                                                   http://host148.ipowerweb.com/~nywolfor
                                                                               •
with the right management              more information on
plans and some                         protecting the predator
                                                                               Conservation Northwest
considerations for residents           species in your state, please       •
living in or near these                review the list of conservation             http://www.conservationnw.org
                                                                               •
predator habitats, both                groups’ website included to
humans and predators can               the right. Each offer many
                                                                               International Wolf Center
                                                                           •
live in harmony.                       ways you can help in the
                                                                                   http://www.wolf.org
                                       plight to rescue, restore, and          •
                                       maintain top predators in the
To prevent ecosystems all              United States and                       Defenders of Wildlife
                                                                           •
                                       internationally.
over North America from                                                            http://www.defenders.org
                                                                               •
experiencing convulsions
brought about by trophic
cascades, the full spectrum of
ecological processes that
operates to perpetuate
biodiversity, especially
predation, must be widely
maintained.
                    Terborgh, et al.


    You can take part in their
valiant efforts to save and
protect these top predators.
With your vote, your time,
your contributions, your
consideration, you can help
maintain biodiversity in the
ecosystems across the
United States.

Article composed by Stacia Clayton. Materials derived from “The Role of
Top Carnivores in Regulating Terrestrial Ecosystem” by Terborgh, John;
Estes, James; Paquet, Paul; Ralls, Katherine; Boyd-Heger, Diane; Miller,
Brian; and Noss, Reed.

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Environmental Writing Final Rewrite

  • 1. Predators. . . Around the world, predators are practicing survival of the fittest. This surviving has been in action for more than a million years for most species. However, extinction rates post human- introduction have multiplied by the hundreds, possibly thousands. This question, then, is posed: Why are so many species now disappearing at an accelerated pace? The sleek, sheen coat of the With the incredible shrinking ecosystem. To understand why cougar might for a moment populations of predator species, each element is so crucial to distract you from the glistening scientists find them difficult to biodiversity, let’s recall the food canines capable of ripping flesh study. These predators are chain. from bone. Sunning at the zoo, reduced to fractions of their Every living organism in an these ‘big cats’ hardly appear to former habitats and are ecosystem supports other living retain their natural predatory instinctively secretive; it is not organisms. For example, in Figure ways. Here, safely locked away surprising that their sightings are 1 the vegetation supports the behind thick glass walls, they rare. Without ample research rabbit. The stimulate our curiosity, our subjects, scientists are quick to rabbit in interest, even perhaps our desire. dismiss studies on predators. turn One can be captivated by the However, in studying their supports power of the majestic, fluid absence, we can still draw very the wolf. movements of these magnificent important conclusions about these The wolf, creatures. species. after dying, Still, this awe and amazement is turned has done little to aid the cats in into Top predators are essential to surviving the onset of humans. necessary the integrity of ecological The cougar population, much like nutrients in that of the gray wolves, has the soil that communit ies… it is imperative steadily declined with the will support to retain or restore top increase of human populations. the growth Farmers, ranchers, and residents of more predators to as many parts of of these areas seem to have no vegetation. North America as pract ical… complaints with the disappearing This is a predators. That is until the prey minimal Figure 1 Terborgh, et al. these predators had been keeping representation; food chains are in in check trample their gardens, actuality much more complex and eat their crops, and disease their One such conclusion is that the usually compound into a food web herds. loss of top predators world-wide (as each organism relies on more The prevalent eradication of has upset the biodiversity of many than one source for its survival). top predators from ecosystems ecosystems. Biodiversity is the However, as you can see from this has agitated the regulatory balance between all organisms simple depiction, removing one balance these predators once occupying an ecological any single element will interrupt provided. Our removal of community. Varied vegetation, the life-supporting role and the predatory species has eliminated herbivore grazing, and predation balance in which each element their role of regulating prey all play important roles in regulates the other elements. This species’ populations. maintaining biodiversity in each is called a trophic cascade.
  • 2. We can find trophic cascades cottonwoods along the herbivores onto predator-free in both top-down and bottom-up streambanks and heavy bank islands by sailors centuries ago to processes at work in ecosystems. erosion…” The same effect has ensure a food supply upon “Top-down” stems from the levels been noted in Yellowstone National successive visits. Without any of the tropic chain (i.e., the food Park (as demonstrated in the video natural predators to control the chain) being manipulated from the linked below) with the removal of number of herbivores, the plant- top of the chain down. To expand wolves and the dramatic increase eaters destroyed the vegetation of on this, return to our previous in the number of elks. the islands. On islands like example. Removing the wolf from The amazing factor not yet Ascension, Juan Fernandez, the the simple food chain would allow considered is how these trophic California Channel Islands, and St. the rabbit to overpopulate (as cascades work in detriment to the George Island, the herbivores’ rabbits are known to do). As a species whose numbers explode. destruction of the existing plant- result, the vegetation the rabbit They quickly diminish their own life lead to their own downfall, as population survives on would resources, resources that also they now had no resources on suffer, unable to keep up with the support other species. To explain, which to survive. increase of its superior trophic think of the food supply that both white-tailed deer and rabbits rely Mammals that would have been level’s increase. The opposite of part of the prey pool of missing this effect is true in bottom-up on. With the elimination of the processes. If we were to increase white-tailed deer’s main carnivores, such as wolves and the number of plants available, this predator, the deer will begin to cougars, have become notoriously too could cause the rabbit consume more of the food supply. Rabbits, in turn, are then abundant to the point that some of population, and thus the wolf population, to amplify. left with less food and their them are now nuisances: by being Trophic cascades are being population decreases. This cause-and-effect action is known road hazards, by browsing recognized in national parks across America; one such example as the “Paine effect.” Through ornamental shrubbery, by raiding has been documented in The the Paine effect, we can trashcans, by preying on birds and Journal of Biological Conservation, determine that removing the top predator of an ecosystem results their nests, by destroying vegetable remarking on the loss of cougars in Zion National Park. Professor in loss of diversity of the prey gardens, and by flooding people’s Robert Beschta of Oregon State community; again, we can yards. University states, “When park conclude that top predators are Terborgh, et al. development caused cougars to absolutely crucial to maintaining begin leaving Zion Canyon in the biodiversity (which is essential 1930s, it allowed much higher for all species’ survival). In the modern era, humans are levels of deer browsing. That set in Reviewing multiple ecosystems responsible for the missing top motion a long cascade of changes around the globe supports this predators. In their absence, we that resulted in the loss of most conclusion. One such example was are noticing an explosion of their More on Trophic Cascades The standard definition of trophic is “of or relating to nutrition.” Each level of the food chain is a trophic level. A trophic cascade is where different levels of the food chain or web are affected by the loss or addition of another element. To see exactly how trophic cascades work, visit: http://youtube.com/watch? v=O8BV0VCn1JU for a short video.
  • 3. former prey. Prey species’ caribou, elk, deer, and moose populations expanding have populations have declined. caused a noticeable amount While these examples have not of problems for humans. been controlled experiments Deer-related car accidents and not always carefully number into the thousands monitored, their results are annually; moose, too, stray consistent. Human interference into highways at their active in nature’s design, specifically times, which happens after the intricate roles of the trophic dark when drivers’ visibility is ladder, typically produces already reduced. Raccoons negative reactions throughout and opossums are just as the ecosystem for all species responsible for overturned involved. trashcans as stray dogs; The conclusion that top beavers, misplaced from their natural territories and without predators play a major predators to keep their regulatory role seems numbers in check, build dams inescapable. in the most inconvenient places flooding residential Terborgh, et al. yards. As frustrating as each of these situations may be, As more and more people have no where to environmental issues appear place the blame outside of on the screens of our our own species. Our televisions, the front pages of reduction of natural habitats our newspapers, and in the and elimination of natural headlines of our favorite online predators has led to the news, we cannot escape the complications we are now importance of conserving experiencing, all a product of nature. A top concern for the loss of biodiversity. conservationists is maintaining We have also been known biodiversity; one guaranteed to upset biodiversity by way to destabilize an introducing predators into ecosystem is by removing the areas previously devoid of top predators from it. Their predators. When the brown removal causes a chain tree snake was introduced reaction; first, we see an and took up residence in overabundance of targeted Guam, it flourished. prey. This consequently affects Consequently, the native bird other species who consume the population drastically same food as the now decreased. With the overabundant species. It also reintroduction of the gray disrupts and destroys wolf in the northern Midwest vegetation and habitats in the area of the United States, ecosystem. There is hope... More on Ecosystems An ecosystem is an interconnected community of living organisms and the physical environment in which they interact to form a life-support system. You don’t have to travel to Yellowstone or a state park to witness an ecosystem at work; you can see one in your own backyard! Discovery Channel’s Animal Planet hosts a show, Backyard Habitat, which assists families with developing ecosystems at their homes. Visit: http://animal.discovery.com/ fansites/backyard/backyard.html for example habitats, tips on renovating your own yard, or to see how to have your yard certified through the National Wildlife Federation.
  • 4. For more information: Despite the gray wolf’s The answer to the opening protected status as an question remains without a Endangered Species and the single, conclusive answer. Keystone Conservation • proven need for the cougar in Many factors contribute to the http://www.keystoneconservation.us • maintaining balance within loss of species. However, ecosystems, both are legally some factors we can control. hunted within several states We must take direct action to Center for Biological Diversity • (Cougars are currently not on restoring and maintaining top http://www.biologicaldiversity.org • the Threatened or predators because their Endangered Species lists; irreplaceable and critical Mountain Lion Foundation however, their sightings are roles inside ecosystems • so rare there is no national or affects practically every other http://www.mountainlion.org • state-specific count of the organism dependant upon that ecosystem. We are part remaining cats in the wild). Wildlife Rescue & Rehabilitation, Inc. • of their ecosystems. At this time, though, there are http://www.wildlife-rescue.org groups working to limit and • hopefully, eventually remove these species from open- In addition to visiting your Wolf Conservation Center • game lists. They propose that state’s official website for http://host148.ipowerweb.com/~nywolfor • with the right management more information on plans and some protecting the predator Conservation Northwest considerations for residents species in your state, please • living in or near these review the list of conservation http://www.conservationnw.org • predator habitats, both groups’ website included to humans and predators can the right. Each offer many International Wolf Center • live in harmony. ways you can help in the http://www.wolf.org plight to rescue, restore, and • maintain top predators in the To prevent ecosystems all United States and Defenders of Wildlife • internationally. over North America from http://www.defenders.org • experiencing convulsions brought about by trophic cascades, the full spectrum of ecological processes that operates to perpetuate biodiversity, especially predation, must be widely maintained. Terborgh, et al. You can take part in their valiant efforts to save and protect these top predators. With your vote, your time, your contributions, your consideration, you can help maintain biodiversity in the ecosystems across the United States. Article composed by Stacia Clayton. Materials derived from “The Role of Top Carnivores in Regulating Terrestrial Ecosystem” by Terborgh, John; Estes, James; Paquet, Paul; Ralls, Katherine; Boyd-Heger, Diane; Miller, Brian; and Noss, Reed.