Primitive vascular plants evolved adaptations for life on land including vascular tissue for transport and alternation of generations. They include psilophytes which were the earliest vascular plants with no specialized organs, lycophytes with true roots, stems and scale-like leaves requiring water for reproduction, and sphenophytes with whorled leaves and siliceous stems requiring water for motile sperm. Advanced ferns have well-developed vascular tissue and xylem and phloem, thriving in damp habitats with a dominant sporophyte and inconspicuous gametophyte reproducing via spores in sori on fronds.