5. . introduction 5
Yeates & Irwin 1996 – study of Apioceridae relationships
Apioceridae Apiocera (Pyrocera) western North America area relationships
Apiocera (Ripidosyrma) South Africa
Apiocera (Anypenus) Argentina / Chile
Apiocera (Apiocera) Australia
6. . introduction 6
Yeates & Irwin 1996 – study of Apioceridae relationships
Apioceridae Apiocera (Pyrocera) western North America area relationships
Mydidae Rhaphiomidas
Apiocera (Ripidosyrma) South Africa
Tongamya
Apiocera (Anypenus) Argentina / Chile
Megascelus
Neorhaphiomidas
Apiocera (Apiocera) Australia
remaining Mydidae
7. .. phylogenetic analysis 7
Taxon Sampling
◊ 134 Mydidae species (= 28.4 %)
› 46 genera (= 69.7 %)
› 7 tribal taxa (= 87.5 %)
› 11 subfamily taxa (= 100.0 %)
◊ 27 Apiocera species (= 19.6 %) Mydidae
◊ Outgroup taxa
› 19 Asilidae species
› 7 Asiloidea species
› 3 Nemestrinoidea
› 1 Tabanomorpha
Apioceridae
collecting localities of specimens included
8. .. phylogenetic analysis 8
Characters
◊ 214 discrete, parsimony informative characters
◊ 36 non-genitalic features sexually dimorphic
tagma # characters remarks
Head 44 10 sexually dimorphic
Thorax 103 22 sexually dimorphic
47 chaetotaxy, sclerites etc.
23 wing
33 leg
Abdomen 67 4 sexually dimorphic
11 chaetotaxy, sclerites etc.
30 ? terminalia
26 / terminalia
◊ DNA data are being gathered
10. .. phylogenetic analysis 10
Phylogenetic analysis
◊ TNT 1.1 (July 2012) Goloboff et al. 2008
◊ exhaustive search of tree space (MacBook Pro 2.2 GHz Intel Core i7)
command(s) time (min) # re-arrangements examined
mult300 01:04 8,383,595,488
xmult=rss hits 20 fuse 2 drift 3 ratchet 10 multiply consense 10 12:31 79,691,144,513
random sectorial searches after hitting best length, find additional trees
by fusing suboptimal with optimal trees
hit best length 20x
consense until strict consensus is stabilized 10x
◊ Bremer support Bremer 1988
◊ DIVA 1.2 Ronquist 1994, Nylander Mac version
11. ... results 11
Results Limata tenuicornis F
Eulonchus sp Six Rivers M
Neorhynchocephalus volaticus
Nemestrinus nigrovillosus
Bombylius major
Dipalta serpentina
Toxophora maxima
Prorates sp Escalante
Apsilocephala longistyla
Hemigephyra atra
◊ 440 most parsimonious trees
Phycus frommeri
Asilidae
Diogmites grossus
Ospriocerus aeacus
Gonioscelis ventralis
Emphysomera pallidapex
Ommatius tibialis
Neolophonotus bimaculatus
Machimus occidentalis
Philonicus albiceps
Hoplistomerus nobilis
Pilica formidolosa
Laphria aktis
Dioctria hyalipennis
Pegesimallus laticornis
Leptogaster cylindrica
Euscelidia pulchra
Stichopogon punctum
◊ 1837 steps
Lasiopogon cinctus
Damalis monochaetes
Holcocephala calva
Apiocera Ap imminuta
Apioceridae
Apiocera Ap tonnoiri
Apiocera R alastor M
Apiocera R braunsi
Apiocera P bilineata
Apiocera P mexicana M
Apiocera P barri
Apiocera P martinorum
Apiocera P parahydra
Apiocera P haruspex
Apiocera P aldrichi
Apiocera P painteri
Apiocera P arena
Apiocera P parkeri
Apiocera P sp Colnett M
◊ strict consensus
Apiocera An obscura
Apiocera An philippii
Apiocera Ap sp Wurarga
Apiocera Ap asilica
Apiocera Ap brevicornis
Apiocera Ap sp Stuart M
Apiocera Ap hardyi
Apiocera Ap pica
Apiocera Ap sp Lloyd
Apiocera Ap similis
Apiocera Ap maritima
Apiocera Ap minor
› several polytomies
Apiocera Ap newmani
Rhaphiomidas hasbroucki
Rhaphiomidas maehleri
Mydidae
Rhaphiomidas parkeri
Tongamya miranda
Megascelus albovillosus
Megascelus melanoproctus
Megascelus nigricornis
Neorhaphiomidas sp Mingenew M
Neorhaphiomidas sp Yanchep
Neorhaphiomidas hardyi
Neorhaphiomidas sp Mandurah
› not well-resolved are:
Cretomydas santanensis
Diochlistus auripennis M
Diochlistus gracilis
Diochlistus melleipennis
Anomalomydas mackerrasi
Anomalomydas australicus F
Midacritus stuardoanus M
Midacritus wagenknechti
Apiophora paulseni
Apiophora quadricinctata
Miltinus stenogaster
Miltinus viduatus
Miltinus maculipennis
› earliest branchings within Apioceridae
Miltinus minutus
Ectyphus armipes
Parectyphus namibiensis
Ectyphus amboseli M
Ectyphus pinguis
Heteromydas bicolor
Opomydas townsendi
Paramydas igniticornis
Plyomydas peruviensis M
Plyomydas sp nov Tandil
Dolichogaster brevicornis
Messiasia californica M
› derived Mydidae ingroup
Messiasia painteri
Mydas clavatus
Mydas tricinctus
Mydas xanthopterus
Stratiomydas lividus
Cacatuopyga basifascia
Cacatuopyga ruficornis M
Phyllomydas bruesii
Phyllomydas parvulus
Pseudorhopalia mirandai
Rhopalia annulata
Rhopalia cincta
Rhopalia olivieri
Perissocerus dumonti F
Perissocerus transcaspicus
Hessemydas parkeri M
Hessemydas daugeroni
Hessemydas seyrigi
Halterorchis sp nov Cape Recife M
Haplomydas crassipes
Eremohaplomydas desertorum
Lachnocorynus sp nov Kotwa M
Leptomydas notos
Nemomydas pantherinus
Nemomydas melanopogon
Namadytes cimbebasiensis
Namadytes vansoni
Nemomydas brachyrhynchus
Nemomydas venosus
Leptomydas lusitanicus
Eremomidas bek
Leptomydas rapti M
Leptomydas turcicus
Leptomydas corsicanus
Nemomydas gruenbergi M
Leptomydas sardous
Nemomydas lamia
Afroleptomydas A inhacae
Afroleptomydas A paganus
Afroleptomydas A suffusipennis
Afroleptomydas C sp Gobabeb
Hispanomydas hispanicus
Mydaselpis ngurumani
Nomoneura caffra
Nomoneura paradoxa
Nomoneuroides natalensis
Syllegomydas S algiricus
Limata tenuicornis F
Eulonchus sp Six Rivers M Syllegomydas S lineatus
Syllegomydas S palestinensis
Neorhynchocephalus volaticus
Nemestrinus nigrovillosus
Bombylius major
Dipalta serpentina
Syllegomydas S proximus
Toxophora maxima
Prorates sp Escalante
Apsilocephala longistyla
Hemigephyra atra
Syllegomydas S vittatus
Phycus frommeri
Diogmites grossus
Ospriocerus aeacus
Syllegomydas sp nov Ruacana
Gonioscelis ventralis
Emphysomera pallidapex
Ommatius tibialis
Neolophonotus bimaculatus
Stenomydas gen nov sp nov Panne M
Machimus occidentalis
Philonicus albiceps
Hoplistomerus nobilis
Pilica formidolosa
Pseudonomoneura hirta
Laphria aktis
Dioctria hyalipennis
Pegesimallus laticornis
Leptogaster cylindrica
Pseudonomoneura micheneri
Euscelidia pulchra
Stichopogon punctum
Lasiopogon cinctus
Afroleptomydas C mauricei
Damalis monochaetes
Holcocephala calva
Apiocera Ap imminuta
Apiocera Ap tonnoiri
Afroleptomydas C sp Stampriet
Apiocera R alastor M
Apiocera R braunsi
Apiocera P bilineata
Apiocera P mexicana M
Pseudonomoneura bajaensis
Apiocera P barri
Apiocera P martinorum
Apiocera P parahydra
Eremomidas sp nov Tiega M
Apiocera P haruspex
Apiocera P aldrichi
Apiocera P painteri
Apiocera P arena
Eremomidas arabicus
Apiocera P parkeri
Apiocera P sp Colnett M
Apiocera An obscura
Apiocera An philippii
Halterorchis karooensis F
Apiocera Ap sp Wurarga
Apiocera Ap asilica
Apiocera Ap brevicornis
Apiocera Ap sp Stuart M
Neolaparopsis puncturatus
Apiocera Ap hardyi
Apiocera Ap pica
Apiocera Ap sp Lloyd
Mydaselpis sp nov Daan
Apiocera Ap similis
Apiocera Ap maritima
Apiocera Ap minor
Apiocera Ap newmani
Mydaselpis sp nov Gamsberg M
Rhaphiomidas hasbroucki
Rhaphiomidas maehleri
Rhaphiomidas parkeri
Tongamya miranda
Vespiodes leopolditertii
Megascelus albovillosus
Megascelus melanoproctus
Megascelus nigricornis
Vespiodes phaios
Neorhaphiomidas sp Mingenew M
Neorhaphiomidas sp Yanchep
Neorhaphiomidas hardyi
Neorhaphiomidas sp Mandurah
Afroleptomydas A consanguinensis
Cretomydas santanensis
Diochlistus auripennis M
Diochlistus gracilis
Diochlistus melleipennis
Afroleptomydas A opacicinctus
Anomalomydas mackerrasi
Anomalomydas australicus F
Midacritus stuardoanus M
Afroleptomydas A sp Clanwilliam
Midacritus wagenknechti
Apiophora paulseni
Apiophora quadricinctata
Miltinus stenogaster
Afroleptomydas A sp Obib
Miltinus viduatus
Miltinus maculipennis
Miltinus minutus
Ectyphus armipes
Afroleptomydas C sp Koeroegab
Parectyphus namibiensis
Ectyphus amboseli M
Ectyphus pinguis
Heteromydas bicolor
Afroleptomydas A omeri
Opomydas townsendi
Paramydas igniticornis
Plyomydas peruviensis M
Afroleptomydas A rufithorax
Plyomydas sp nov Tandil
Dolichogaster brevicornis
Messiasia californica M
Messiasia painteri
Arenomydas partitus
Mydas clavatus
Mydas tricinctus
Mydas xanthopterus
Stratiomydas lividus
Arenomydas caerulescens
Cacatuopyga basifascia
Cacatuopyga ruficornis M
Phyllomydas bruesii
Arenomydas callosus
Phyllomydas parvulus
Pseudorhopalia mirandai
Rhopalia annulata
Rhopalia cincta
Arenomydas namaquensis M
Rhopalia olivieri
Perissocerus dumonti F
Perissocerus transcaspicus
Hessemydas parkeri M
Notosyllegomydas brincki
strict consensus topology (2121 steps)
Hessemydas daugeroni
Hessemydas seyrigi
Halterorchis sp nov Cape Recife M
Haplomydas crassipes
Eremohaplomydas desertorum
Lachnocorynus sp nov Kotwa M
Leptomydas notos
Syll gen nov sp nov Gobabeb
Cephalocera longirostris
Nemomydas pantherinus
Nemomydas melanopogon
Namadytes cimbebasiensis
Namadytes vansoni
Cephalocera sp Papendorp
Nemomydas brachyrhynchus
Nemomydas venosus
Leptomydas lusitanicus
Eremomidas bek
Cephalocera sp Sors Sors
Leptomydas rapti M
Leptomydas turcicus
Leptomydas corsicanus
Cephalocera imitata M
Nemomydas gruenbergi M
Leptomydas sardous
Nemomydas lamia
Afroleptomydas A inhacae
Cephalocera sp Ratelfontein
Afroleptomydas A paganus
Afroleptomydas A suffusipennis
Afroleptomydas C sp Gobabeb
Hispanomydas hispanicus
Syllegomydas S elachys
Mydaselpis ngurumani
Nomoneura caffra
Nomoneura paradoxa
Syllegomydas N astrictus
Nomoneuroides natalensis
Bremer ≥ 3
Syllegomydas S algiricus
Syllegomydas S lineatus
Syllegomydas S palestinensis
Namibimydas psamminos
Syllegomydas S proximus
Syllegomydas S vittatus
Syllegomydas sp nov Ruacana
Stenomydas gen nov sp nov Panne M
Namibimydas stuckenbergi
Pseudonomoneura hirta
Pseudonomoneura micheneri
Afroleptomydas C mauricei
Afroleptomydas C sp Stampriet
Nothomydas picketti
Pseudonomoneura bajaensis
Eremomidas sp nov Tiega M
Eremomidas arabicus
Cleamydas gen nov sp nov Ugab F
Halterorchis karooensis F
Neolaparopsis puncturatus
Mydaselpis sp nov Daan
Mydaselpis sp nov Gamsberg M
Syll gen nov sp nov minute proboscis F
Vespiodes leopolditertii
Vespiodes phaios
Afroleptomydas A consanguinensis
Afroleptomydas A opacicinctus
Syllegomydas N arnoldi M
Afroleptomydas A sp Clanwilliam
Afroleptomydas A sp Obib
Afroleptomydas C sp Koeroegab
Syllegomydas N dispar
Afroleptomydas A omeri
Afroleptomydas A rufithorax
Arenomydas partitus
Arenomydas caerulescens
Syllegomydas N heothinos
Arenomydas callosus
Arenomydas namaquensis M
Notosyllegomydas brincki
Syll gen nov sp nov Gobabeb
Diochlistinae Argentina F
Cephalocera longirostris
Cephalocera sp Papendorp
Cephalocera sp Sors Sors
Cephalocera imitata M
Cephalocera sp Ratelfontein
Syllegomydas S elachys
Syllegomydas N astrictus
Mitrodetus nanoglossa M
Mitrodetus microglossa
Namibimydas psamminos
Namibimydas stuckenbergi
Nothomydas picketti
Cleamydas gen nov sp nov Ugab F
Mitrodetus dentitarsis
Syll gen nov sp nov minute proboscis F
Syllegomydas N arnoldi M
Syllegomydas N dispar
Syllegomydas N heothinos
Mitrodetus dimidiatus
Diochlistinae Argentina F
Mitrodetus nanoglossa M
Mitrodetus microglossa
Mitrodetus dentitarsis
Mitrodetus dimidiatus
12. ... results 12
Placement of † Cretomydas santanensis Mydidae
Rhaphiomidas hasbroucki
Rhaphiomidas maehleri
Rhaphiomidas parkeri
Tongamya miranda
Megascelus albovillosus
Megascelus melanoproctus
Megascelus nigricornis
◊ explicitly included in matrix
Neorhaphiomidas hardyi
≈ 112 myo
Neorhaphiomidas sp Mandurah
Neorhaphiomidas sp Mingenew M
Neorhaphiomidas sp Yanchep
Cretomydas santanensis
Diochlistus auripennis M
Diochlistus gracilis
Diochlistus melleipennis
Anomalomydas mackerrasi
Anomalomydas australicus F
Midacritus stuardoanus M
Midacritus wagenknechti
◊ 80% missing data
Apiophora paulseni
Apiophora quadricinctata
Miltinus stenogaster
Miltinus viduatus
Miltinus maculipennis
Miltinus minutus
Ectyphus armipes
Ectyphus amboseli M
Ectyphus pinguis
Parectyphus namibiensis
Heteromydas bicolor
Opomydas townsendi
◊ wing venation very similar to Diochlistus
Paramydas igniticornis
Messiasia californica M
Messiasia painteri
Plyomydas sp nov Tandil
Plyomydas peruviensis M
Cacatuopyga basifascia
Cacatuopyga ruficornis M
Mydas clavatus
Mydas tricinctus
(Australia) and Rhaphiomidas (wNA)
Mydas xanthopterus
Stratiomydas lividus
Dolichogaster brevicornis
Phyllomydas bruesii
Phyllomydas parvulus
Pseudorhopalia mirandai
Rhopalia cincta
Rhopalia annulata
Rhopalia olivieri
Perissocerus dumonti F
Perissocerus transcaspicus
Hessemydas parkeri M
Hessemydas daugeroni
Hessemydas seyrigi
Halterorchis sp nov Cape Recife M
Haplomydas crassipes
Eremohaplomydas desertorum
Lachnocorynus sp nov Kotwa M
Leptomydas notos
Nemomydas pantherinus
Nemomydas melanopogon
Namadytes cimbebasiensis
Namadytes vansoni
Nemomydas brachyrhynchus
Nemomydas venosus
Leptomydas lusitanicus
Eremomidas bek
Leptomydas rapti M
Leptomydas turcicus
Leptomydas corsicanus
Nemomydas gruenbergi M
Leptomydas sardous
Pseudonomoneura hirta
Pseudonomoneura micheneri
Pseudonomoneura bajaensis
Eremomidas sp nov Tiega M
Eremomidas arabicus
Halterorchis karooensis F
Nemomydas lamia
Arenomydas partitus
Arenomydas caerulescens
Arenomydas callosus
Arenomydas namaquensis M
Notosyllegomydas brincki
Syll gen nov sp nov Gobabeb
Cephalocera longirostris
Cephalocera sp Papendorp
Cephalocera sp Sors Sors
Cephalocera imitata M
Cephalocera sp Ratelfontein
Mydaselpis ngurumani
Neolaparopsis puncturatus
Mydaselpis sp nov Daan
Mydaselpis sp nov Gamsberg M
Vespiodes leopolditertii
Vespiodes phaios
Nomoneura caffra
Nomoneura paradoxa
Limata tenuicornis F
Eulonchus sp Six Rivers M
Nomoneuroides natalensis
Afroleptomydas A suffusipennis
Neorhynchocephalus volaticus
Nemestrinus nigrovillosus
Bombylius major
Dipalta serpentina
Toxophora maxima
Afroleptomydas A paganus
Prorates sp Escalante
Apsilocephala longistyla
Hemigephyra atra
Phycus frommeri
Diogmites grossus
Ospriocerus aeacus
Afroleptomydas A inhacae
Gonioscelis ventralis
Emphysomera pallidapex
Ommatius tibialis
Neolophonotus bimaculatus
Machimus occidentalis
Afroleptomydas A consanguinensis
Philonicus albiceps
Hoplistomerus nobilis
Pilica formidolosa
Laphria aktis
Dioctria hyalipennis
Afroleptomydas A opacicinctus
Pegesimallus laticornis
Leptogaster cylindrica
Euscelidia pulchra
Stichopogon punctum
Lasiopogon cinctus
Damalis monochaetes
Afroleptomydas A sp Clanwilliam
Holcocephala calva
Apiocera Ap imminuta
Apiocera Ap tonnoiri
Apiocera R alastor M
Apiocera R braunsi
Afroleptomydas C sp Koeroegab
Apiocera P bilineata
Apiocera P mexicana M
Apiocera P barri
Apiocera P martinorum
Apiocera P parahydra
Afroleptomydas A sp Obib
Apiocera P haruspex
Apiocera P aldrichi
Apiocera P painteri
Apiocera P arena
Apiocera P parkeri
Apiocera P sp Colnett M
Afroleptomydas A omeri
Apiocera An obscura
Apiocera An philippii
Apiocera Ap sp Wurarga
† Cretomydas santanensis, Crato Formation, Brazil
Apiocera Ap asilica
Apiocera Ap brevicornis
Afroleptomydas A rufithorax
Apiocera Ap sp Stuart M
Apiocera Ap hardyi
Apiocera Ap pica
Apiocera Ap sp Lloyd
Apiocera Ap similis
Afroleptomydas C mauricei
Apiocera Ap maritima
Apiocera Ap minor
Apiocera Ap newmani
Rhaphiomidas hasbroucki
Rhaphiomidas maehleri
Rhaphiomidas parkeri
Afroleptomydas C sp Stampriet
Tongamya miranda
Megascelus albovillosus
Megascelus melanoproctus
Megascelus nigricornis
Neorhaphiomidas sp Mingenew M
Afroleptomydas C sp Gobabeb
Neorhaphiomidas sp Yanchep
Neorhaphiomidas hardyi
Neorhaphiomidas sp Mandurah
Cretomydas santanensis
Diochlistus auripennis M
Stenomydas gen nov sp nov Panne M
Diochlistus gracilis
Diochlistus melleipennis
Anomalomydas mackerrasi
Anomalomydas australicus F
Midacritus stuardoanus M
Midacritus wagenknechti
Syllegomydas S lineatus
Apiophora paulseni
Apiophora quadricinctata
Miltinus stenogaster
Miltinus viduatus
Miltinus maculipennis
Syllegomydas sp nov Ruacana
Miltinus minutus
Ectyphus armipes
Parectyphus namibiensis
Ectyphus amboseli M
Ectyphus pinguis
Hispanomydas hispanicus
Heteromydas bicolor
Opomydas townsendi
Paramydas igniticornis
Plyomydas peruviensis M
Plyomydas sp nov Tandil
Syllegomydas S vittatus
Dolichogaster brevicornis
Messiasia californica M
Messiasia painteri
Mydas clavatus
Mydas tricinctus
Mydas xanthopterus
Syllegomydas S palestinensis
Stratiomydas lividus
Cacatuopyga basifascia
Cacatuopyga ruficornis M
Phyllomydas bruesii
Phyllomydas parvulus
Syllegomydas S algiricus
Pseudorhopalia mirandai
Rhopalia annulata
Rhopalia cincta
Rhopalia olivieri
Perissocerus dumonti F
Syllegomydas S proximus
Perissocerus transcaspicus
Hessemydas parkeri M
Hessemydas daugeroni
Hessemydas seyrigi
Halterorchis sp nov Cape Recife M
Haplomydas crassipes
Syllegomydas S elachys
Eremohaplomydas desertorum
Lachnocorynus sp nov Kotwa M
Leptomydas notos
Nemomydas pantherinus
Nemomydas melanopogon
Syllegomydas N astrictus
Namadytes cimbebasiensis
Namadytes vansoni
Nemomydas brachyrhynchus
Nemomydas venosus
Leptomydas lusitanicus
Namibimydas psamminos
Eremomidas bek
Leptomydas rapti M
Leptomydas turcicus
1 of the mp trees
Leptomydas corsicanus
Nemomydas gruenbergi M
Leptomydas sardous
Namibimydas stuckenbergi
Nemomydas lamia
Afroleptomydas A inhacae
Afroleptomydas A paganus
Afroleptomydas A suffusipennis
Afroleptomydas C sp Gobabeb
Nothomydas picketti
Hispanomydas hispanicus
Mydaselpis ngurumani
Nomoneura caffra
Nomoneura paradoxa
Nomoneuroides natalensis
Cleamydas gen nov sp nov Ugab F
Syllegomydas S algiricus
Syllegomydas S lineatus
Syllegomydas S palestinensis
Syllegomydas S proximus
Syllegomydas S vittatus
Syllegomydas sp nov Ruacana
Syll gen nov sp nov minute proboscis F
Stenomydas gen nov sp nov Panne M
Pseudonomoneura hirta
Pseudonomoneura micheneri
Afroleptomydas C mauricei
Afroleptomydas C sp Stampriet
Syllegomydas N arnoldi M
Pseudonomoneura bajaensis
Eremomidas sp nov Tiega M
Eremomidas arabicus
Halterorchis karooensis F
Neolaparopsis puncturatus
Syllegomydas N dispar
Mydaselpis sp nov Daan
Mydaselpis sp nov Gamsberg M
Vespiodes leopolditertii
Vespiodes phaios
Afroleptomydas A consanguinensis
Afroleptomydas A opacicinctus
Syllegomydas N heothinos
Afroleptomydas A sp Clanwilliam
Afroleptomydas A sp Obib
Afroleptomydas C sp Koeroegab
Afroleptomydas A omeri
Afroleptomydas A rufithorax
Diochlistinae Argentina F
Arenomydas partitus
Arenomydas caerulescens
Arenomydas callosus
Arenomydas namaquensis M
Notosyllegomydas brincki
Mitrodetus nanoglossa M
Syll gen nov sp nov Gobabeb
Cephalocera longirostris
Cephalocera sp Papendorp
Cephalocera sp Sors Sors
Cephalocera imitata M
Cephalocera sp Ratelfontein
Mitrodetus microglossa
Syllegomydas S elachys
Syllegomydas N astrictus
Namibimydas psamminos
Namibimydas stuckenbergi
Nothomydas picketti
Mitrodetus dentitarsis
Cleamydas gen nov sp nov Ugab F
Syll gen nov sp nov minute proboscis F
Syllegomydas N arnoldi M
Syllegomydas N dispar
Syllegomydas N heothinos
Mitrodetus dimidiatus
Diochlistinae Argentina F
Mitrodetus nanoglossa M
Mitrodetus microglossa
Mitrodetus dentitarsis
Mitrodetus dimidiatus