2. Area: 256.371 Km²
198,985 sq.miles
Population: 13,810,000
Language: Spanish
Capital: Quito
Major City: Guayaquil
3. ECUADOR
La Mitad del Mundo
Puerto Lopez
Salango Salango Island
Guayaquil Downtown Malecon Museum - Guayaquil
4. ECUADOR
Canta la Piedra Puerto Lopez
Shell trash Local child fisherman
Guayaquil Puerto Cayo Fisherman Village Cotopaxa
5. INTERTIDAL SHELLS
EXPOSED ROCKS
AND TIDE POOLS
Nerita funiculata Menke, 1851
13mm
Nerita scabricosta Tegula picta (McLean, 1970) Tegula panamensis
Lamarck, 1822 – 14mm 15mm (Philippi, 1849) - 25mm
Fissurella microtrema Fissurella longifissa Littorina aspera Littorina zebra Donovan, 1825 Thais biserialis Purpura pansa Gould, 1853
Sowerby, 1835 - 22mm Sowerby, 1863 - 30mm Philippi, 1846 - 2mm 33mm (Blainville, 1832) - 36mm 35mm
Fissurella virescens Scurria mesoleuca Morula lugubris Conus nux Broderip, 1833 Acanthina brevidentata Chiton stokesii Broderip, 1833
Sowerby, 1835 - 30mm (Menke, 1851) - 20mm (C.B.Adams, 1852) - 8mm 13mm (Wood, 1828) - 17mm 52mm
6. INTERTIDAL
SHELLS
ROCKS AND SAND BEACH Anachis rugulosa Anachis strongi Bartsch, 1928
(Sowerby, 1844) - 10mm 4mm
Anachis pardalis Anachis rugosa Anachis pygmaea Olivella columellaris Oliva undatella ecuatoriana Petuch & Sargent, 1986 - 16mm
(Hinds, 1843) - 6mm (Sowerby, 1832) - 14mm (Sowerby, 1832) - 6mm (Sowerby, 1825) - 10mm
Cerithium gallapaginis Mazatlania fulgurata Donax gracilis Hanley, 1845 Donax mancorensis
Sowerby, 1855 - 8mm (Philippi, 1846) - 13mm 11mm (Olsson, 1961) - 17mm
Donax carinatus Donax peruvianus Donax navicula Hanley, 1845 Hastula cinerea luctuosa Pitar lupanaria (Lesson, 1830) - 42mm
(Hanley, 1843) - 28mm Deshayes, 1855 - 16mm 15mm (Hinds, 1844) - 40mm
7. INTERTIDAL
SHELLS
MANGROVE AND SOFT-ROCKS
Protothaca columbiensis Nassarius pagodus
(Sowerby, 1835) - 48mm (Reeve, 1844) - 22mm
Cerithidea valida Rhinocoryne humboldti Natica unifasciata Lithophaga aristata Petricola denticulata Parapholas calva
(C.B.Adams, 1852) - 38mm (Valenciennes, 1832) - 31mm Lamarck, 1822 - 21mm (Dillwyn, 1817) - 19mm Sowerby, 1834 - 17mm (Sowerby, 1834) - 25mm
Parapholas tubifera Pholadidea esmeraldensis Pholadidea melanura Pholadidea quadra
(Sowerby, 1834) - 25mm (Olsson, 1961) - 18mm (Sowerby, 1834) - 30mm (Sowerby, 1834) - 28mm
Jouannetia duchassaingi Fischer, 1862 - 43mm Parapholas acuminata Jouannetia pectinata Irus ellipticus Clypeolum latissimus
(Sowerby, 1824) - 26mm (Conrad, 1849) - 11mm (Sowerby, 1834) - 18mm fontaineanum (Orbigny, 1840)
20mm
8. SNORKELING SHELLS
ROCK BOTTOM
Cypraea arabicula Cypraea robertsi Cypraea cervinetta Thais melones
Lamarck, 1811 – 30mm (Hidalgo, 1906) – 20mm Kiener, 1843 – 50mm (Duclos, 1832) – 60mm
Thais speciosa Eupleura nitida Turbo saxosus Trivia radians
(Valenciennes, 1832) – 28mm (Broderip, 1833) – 14mm Wood, 1828 – 36mm (Lamarck, 1822) – 16mm
Muricopsis zeteki Hertlein & Trivia sanguinea Bursa caelata Vitularia salebrosa (King & Jenneria pustulata Lightfoot, 1786) – 16mm
Strong, 1951 - 17mm (Sowerby, 1832) – 7mm (Broderip, 1833) - 45 mm Broderip, 1832) – 30mm
9. SNORKELING SHELLS
ROCK AND RUBBLE BOTTOM
Cantharus elegans (Griffith Cantharus ringens
& Pidgeon, 1834) – 46mm (Reeve, 1846) – 21mm
Opeatostoma pseudodon Anachis fluctuata Engina maura Engina tabogaensis
(Burrow, 1815) – 70mm (Sowerby, 1832) - 15mm (Sowerby, 1832) – 18mm Bartsch, 1931 – 12mm
Leucozonia cerata Thais planospira (Lamarck, 1822) – 55mm Anachis varia Columbella fuscata Columbella major
(Wood, 1828) – 54mm (Sowerby, 1832) – 19mm Sowerby, 1832 - 24mm Sowerby, 1832 – 30mm
Latirus hemphilli Hertlein Latirus mediamericanus Anachis fulva Mitrella guttata Columbella haemastoma Columbella strombiformis
& Strong, 1951 - 40mm Hertlein & Strong, 1951 (Sowerby, 1832) – 20mm (Sowerby, 1832) – 11mm Sowerby, 1832 – 23mm Lamarck, 1822 – 23mm
80mm
10. SNORKELING SHELLS
ROCK AND SAND BOTTOM
Mitra lens Mitra tristis Conus brunneus Conus fergusoni Conus princeps lineolatus Valenciennes, 1832 – 33mm
Wood, 1828 – 32mm Broderip, 1836 – 14mm Wood, 1828 – 20mm Sowerby, 1873 – 90mm
Pilsbryspira albinodata Pilsbryspira collaris Modiolus capax Megapitaria squalida chionacea Lima pacifica Barbatia reeveana
(Reeve, 1843) – 22mm (Sowerby, 1834) – 10mm (Conrad, 1837) – 35mm (Menke, 1847) – 33mm Orbigny, 1846 – 25mm (Orbigny, 1876) – 60mm
Periglypta multicostata Protothaca beili Protothaca columbiensis Protothaca metodon (Pilsbry Arca pacifica Arcopsis solida
(Sowerby, 1835) – 85mm (Olsson, 1961) - 50mm (Sowerby, 1835) – 48mm & Lowe, 1932) - 24mm (Sowerby, 1833) – 70mm (Sowerby, 1833) – 14mm
11. DIVING SHELLS
ROCK BOTTOM (5 to 35 meters)
Cypraea nigropunctata Cypraea albuginosa Cypraea aequinoctialis Cymatium gibbosum
Gray, 1828 – 29mm Gray, 1825 - 22mm (Schilder, 1933) – 45mm adairense Dall, 1910 – 32mm
Cymatium vestitum Linatella wiegmanni Turbo mazatlanicus Pilsbry Turbo magnificus Colubraria jordani Colubraria lucasensis Strong
(Hinds, 1844) – 45mm (Anton, 1838) – 85mm & Lowe, 1932 – 9mm Jonas, 1844 – 18mm Strong, 1938 – 35mm & Hertlein, 1937 – 38mm
Vasum caestus Astraea babelis Turbo squamiger Colubraria ochsneri Hertlein Colubraria procera Cantharus sanguinolentus
(Broderip, 1833) – 100mm (Fischer, 1874) – 28mm Reeve, 1843 – 20mm & Allison, 1968 – 25mm (Sowerby, 1832) – 40mm (Duclos, 1833) – 23mm
12. DIVING SHELLS
ROCK BOTTOM (5 to 35 meters)
Engina pulchra Hexaplex radix Homalocantha oxycantha Muricanthus princeps
(Reeve, 1846) – 20mm (Gmelin, 1791) – 80mm (Broderip, 1833) – 70mm (Broderip, 1833) – 75mm
Engina tabogaensis Phyllonotus regius
Bartsch, 1931 – 11mm (Swainson, 1821) – 122mm
Pleuroploca princeps (Sowerby, 1825) – 350mm
Latirus rudis Latirus tumens Cantharus vibex Cantharus lautus Phyllonotus regius (Swainson, 1821) – 130mm
(Reeve, 1847) – 40mm Carpenter, 1856 – 75mm (Broderip, 1833) – 40mm (Reeve, 1846) – 22mm
13. DIVING SHELLS
RUBBLE AND SAND BOTTOM (5 to 35 meters)
Strombus galeatus Strombus peruvianus
Swainson, 1823 – 215mm Swainson, 1823 – 170mm
Strombus granulatus Cyphoma emarginatum Phenacovolva lenoraea Cardin Cypraecassis coarctata Cypraecassis tenuis Malea ringens
Swainson, 1822 – 80mm (Sowerby, 1830) – 20mm & Walls, 1980 – 26mm Sowerby, 1825 – 36mm (Wood, 1828) – 65mm (Swainson, 1822) – 50mm
Americardia guanacastensis Trachycardium pristipleura Conus dalli Conus archon
(Hertlein & Strong, 1947) (Dall, 1901) - 46mm Stearns, 1873 - 60mm Broderip, 1833 – 68mm
Megapitaria aurantiaca Conus lucidus Conus purpurascens Oppenheimopecten perulus Spondylus princeps Broderip, 1833 – 90mm
(Sowerby, 1831) - 55mm Wood, 1828 – 36mm Sowerby, 1833 – 68mm Olsson, 1961 – 35mm
15. DREDGING SHELLS
MUD AND SAND BOTTOM (40 to 100 meters)
Eulima panamensis
(Bartsch, 1917) – 7mm
Niso excolpa Niso interrupta Aesopus arestus
Bartsch, 1917 – 18mm (Sowerby, 1834) – 10mm Dall, 1919 – 7mm
Anachis adelinae Cosmioconcha modesta Leurolepas roseola Epitonium politum Mazatlania hesperia Pilsbry Microcithara uncinata
(Tryon, 1883) – 7mm (Powys, 1835) – 20mm McLean, 1970 – 7mm (Sowerby, 1844) – 9mm & Lowe, 1932 – 10mm (Sowerby, 1832) – 10mm
Strombina gibberula Lucapinella eleanorae Epitonium shyorum DuShane Nassarina perata Nassarina cruentata Nassarina whitei
(Sowerby, 1832) – 10mm McLean, 1967 – 8mm & McLean, 1968 – 7mm Keen, 1971 – 7mm (Mörch, 1860) – 6mm (Bartsch, 1928) – 6mm
16. DREDGING SHELLS
MUD AND SAND BOTTOM (40 to 100 meters)
Oliva radix Petuch & Oliva splendidula
Sargent, 1987 – 40mm Sowerby, 1825 – 33mm
Olivella aureocincta Olivella anazora Olivella fletcherae Olivella guayaquilensis Olivella intorta Phos cumingii
Carpenter, 1857 – 8mm (Duclos, 1835) – 14mm Berry, 1958 – 10mm Bartsch, 1928 – 14mm Carpenter, 1857 – 5mm Reeve, 1846 – 23mm
Drillia clavata Kylix contracta McLean & Cancellaria deroyae Cancellaria mitriformis Cancellaria strongi Cancellaria tessellata
(Sowerby, 1834) – 13mm Poorman, 1971 – 9mm Petit, 1970 – 16mm Sowerby, 1832 – 22mm Shasky, 1961 – 15mm Sowerby, 1832 – 10mm
Perplicaria clarki Compsodrillia jaculum (Pilsbry Compsodrillia thestia Crassispira coracina McLean Trigoniocardia granifera Chione mariae
M.Smith, 1947 – 14mm & Lowe, 1932) – 17mm (Dall, 1919) – 11mm & Poorman, 1971 – 13mm (Broderip et.al., 1829) – 12mm (Orbigny, 1846) – 23mm
17. TRAWLING SHELLS
SAND AND MUDDY BOTTOM (50 to 150 meters)
Solenosteira pallida (Broderip Fusinus dupetitthouarsi Fusinus panamensis Solenosteira anomala
& Sowerby, 1846) – 80mm (Kiener, 1840) – 190mm Dall, 1908 – 160mm (Reeve, 1847) – 45mm
Murex elenensis Pteropurpura centrifuga Distorsio decussata Metula amosi
Dall, 1909 – 75mm (Hinds, 1844) – 85mm (Valenciennes, 1832) – 90mm Vanatta, 1913 – 40mm
Ficus ventricosa Oliva kaleontina Architectonica nobilis Northia pristis (Deshayes in Eupleura muriciformis (Broderip, 1833) – 35mm
(Sowerby, 1825) – 120mm Duclos, 1832 - 32mm Röding, 1798 – 40mm Lamarck, 1844) – 75mm
18. TRAWLING SHELLS
SAND AND MUDDY BOTTOM (50 to 150 meters)
Cancellaria bulbulus Cancellaria cassidiformis Cancellaria indentata Cancellaria decussata
Sowerby, 1832 – 40mm Sowerby, 1832 – 42mm Sowerby, 1832 – 22mm Sowerby, 1832 – 40mm Harpa crenata Swainson, 1822 – 60mm
Trigonostoma bullatum Trigonostoma milleri Conus arcuatus Broderip & Conus fergusoni Conus patricius
(Sowerby, 1832) – 40mm Burch, 1949 – 30mm Sowerby, 1829 – 60mm Sowerby, 1873 – 65mm Hinds, 1843 – 90mm
Subcancilla erythrogramma Conus poormani Conus recurvus Conus regularis Subcancilla hindsii Mitra swainsoni
(Tomlin, 1931) - 32mm Berry, 1960 – 55mm Broderip, 1833 - 75mm Sowerby, 1833 – 60mm (Reeve, 1844) - 42mm Broderip, 1836 – 140mm
20. LAND SHELLS
Aperostoma fischeri Drymaeus hidalgoi
(Hidalgo, 1867) – 44mm (daCosta, 1898) – 40mm
Aperostoma manabense Torre, Porphyrobaphe iostoma (Sowerby, 1824) – 60mm Porphyrobaphe grevillei Isomeria ritchieana Isomeria subcastanea
Bartsch & Morrison, 1942 – 40mm (Sowerby, 1876) – 58mm Pilsbry, 1889 – 60mm (Pfeiffer, 1842) – 38mm
Calaperostoma nigrofasciatum Lagocyclus vasconesi Porphyrobaphe i. bilabratus Porphyrobaphe i. phasianella Lampadion cymatodes Plecochilus semipictus
(Miller, 1879) – 27mm (Jousseaume, 1897) – 32mm Pilsbry, 1899 – 75mm (Valenciennes, 1833) – 60mm (Pfeiffer, 1852) – 42mm (Hidalgo, 1869) – 48mm
21. NEW SPECIES
Phos sp. Compsodrillia sp. Microcithara sp. Pteropurpura sp.
Anachis cf. costellata Anachis cf. teevani Anachis sp. Agladrillia sp.
22. ECUADOR
Number of Species Found
Gastropoda marine 578
land 42
fresh-water 4
Bivalvia marine 218
fresh-water 3
Polyplacophora 6
Scaphopoda 3
Cephalopoda 2
total 856
25. Project & Lay-Out REFERENCE
José Coltro
Abbott, R.T., 1989, Compendium of Landshells
Abbott, R.T. & Dance, S.P., 1983, Compendium of Seashells
Landscape Photography Bequaert, J.C., 1948, Monograph of the Strophocheilidae, a Neotropical Family of
Terrestrial Mollusks , Bull.Mus.Comp.Zoo.Harvard, vol. 100 no1
José Coltro Bratcher, T. & Cernohorsky, W.O.,1987, Living Terebras
Cate, C.N., 1979, A Review of the Triviidae, San Diego Soc.
Cernohorsky, W. O., 1984, Systematics of the Family Nassariidae, Bull.Auckland
Inst.Museum, Number 14
Photography and Art Cernohorsky, W.O., 1991, The Mitridae of the World, part II Ind.Pac.Moll
Carlos Henckes Chaney, H.W., 1987, A Comparaive Study of Two Similar Panamic Cones: Conus
ximenes and Conus mahogani, The Veliger 29(4)
Cossignani, T., 1994, Bursidae of the World
Finet, Y., 1994, Marine Molluscs of the Galapagos, nr.1
Maps Google Earth ® Finet, Y., 1994, The marine mollusks of the Galapagos Islands: a documented fauna list
Fischer-Piette, E., 1977, Revision des Cardiidae, Mem. Mus.Nat. Paris, no.1
Henning, T. & Hemmen, J., 1993, Ranellidae & Personidae of the World
Hickman, C.P.Jr & Finet, Y., 1999, A Field Guide to Marine Molluscs of Galápagos
Kaicher, S.D., Card Catalogue of World-Wide Shells
Keen, M., 1971, Sea Shells of Tropical West America
Kosuge & Suzuki, 1985, Illust.Catal. of Latiaxis and Its Related Groups
Wikipedia ® Kreipl, K., 1997, Recent Cassidae
Lamprell, K., 1987, Spondylus
Lorenz, F. & Fehse, D., 2009, The Living Ovulidae
Lorenz, F. & Hubert, A., 2000, A Guide to Wolrdwide Cowries, 2nd. Edition
Marshall, C.P., 1995, Recent Changes in the Pectinidae part 1, American Conchologist,
vol.23, no.1
Marshall, C.P., 1995, Recent Changes in the Pectinidae part 2, American Conchologist,
vol.23, no.3
Miller, K., 1878-1879, Die Binnenmollusken von Ecuador, Malac.
Parkison, B., Hemmen, J. & Groh, K., 1987, Tropical Landshells of the World
Petuch, E.J. & Sargent, D.M., 1986, Atlas of the Living Oliva
Pilsbry, H.A., 1895-1907, Manual of Conchology, Vol.V, Vol.X, Vol.XI, Vol.XII, Vol.XVIII.
Pilsbry, H. A., 1939, South American Land & Freshwater Mollusks, X—Species of
Colombia & Ecuador, Not.Nat.19
Röckel, D., Korn, W. & Kohn, A.J., 1995, Manual of the Living Conidae, Vol.1
Roth, Barry, 1978, New species and records of tropical west American Marginellidae
(Mollusca: Neogastropoda). Natural History Museum of Los Angeles, Contribution in
Science 292:1-17; figs. 1-16
Simone, L.R.L., 2006, Land and Freshwater Molluscs of Brasil
Skoglund, C. 1992, Additions to the Panamic Province Gastropod (Mollusca) Litterature
1971 to 1992, The Festivus, Vol.XXIV, Supplement
Torre, C.; Bartsch, P. & Morrison, J.P.E., 1942, The cyclophorid operdulate land
mollusks of America. Bull. U.S.Nat.Mus., 181:1-306, 42pls
Turner, R.D., 1955, The Family Pholadidae in the W.Atlantic and E.Pacific, Part II,
Pholadinae, Johnsonia vol.3, no.34
Copyright © Femorale 2009 Weil, A.; Brown., L. and Neville, B., 1999, The Wentletrap Book - Guide to the Recent
Epitoniidae of the World
