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PRIMARY AND
SECONDARY GROWTH
     IN PLANTS
Meristems generate cells for new organs

                                                   Primary growth in stems
Shoot apical
meristems                                                              Epidermis
(in buds)
                                                                             Cortex
                                                                          Primary phloem

                                                                        Primary xylem
                Vascular
                cambium    Lateral                            Pith
                Cork       meristems
                cambium
                                                 Secondary growth in stems
                                                                             Periderm
                                                                               Cork
                                            Pith                               cambium
                                                                                Cortex
                                       Primary                                  Primary
                                       xylem                                    phloem

  Root apical                          Secondary
  meristems                            xylem                                  Secondary
                                                                              phloem
                                                       Vascular cambium
Woody plants
                                                    Terminal bud
                                                    Bud scale

•In woody                                              Axillary buds

plants, primary                                        Leaf scar
                   This year’s growth                              Node
and secondary      (one year old)                     Stem

growth occur                                         Internode

simultaneously                                                           One-year-old side
                                                                         branch formed
but in different                                                         from axillary bud
                                                                         near shoot apex
locations          Last year’s growth
                                                                       Leaf scar
                   (two years old)
                                                                       Scars left by terminal
                                                                       bud scales of previous
                                                                       winters

                     Growth of two      Leaf scar
                     years ago (three
                     years old)
Primary Growth of Roots

                                Cortex   Vascular cylinder
                     Epidermis

Key                                               Zone of
                     Root hair                    maturation
 Dermal
 Ground
 Vascular



                                                   Zone of
                                                   elongation




                     Apical                      Zone of cell
                     meristem                    division

                     Root cap


            100 µm
Primary Growth of Shoots
                  Apical meristem   Leaf primordia


•A shoot
apical
meristem
gives rise to a
repetition of                                        Developing
                                                     vascular
internodes                                           strand
and leaf-
bearing nodes

                                                     Axillary bud
                                                     meristems



                                          0.25 mm
How a plant grows from a seed –
        primary growth




  • fuwopabecu.comoj.com
Secondary growth adds girth to stems
       and roots in woody plants
•Occurs in stems          Primary and secondary growth
                          in a two-year-old stem


and roots of
                   Epidermis

woody plants       Cortex
                   Primary
                   phloem
                                                               Pith
                                                                  Primary xylem
                                                                    Vascular cambium

but rarely in      Vascular
                   cambium
                   Primary
                                                                        Primary phloem
                                                                                    Cortex
                                                                                             Epidermis
                   xylem                                     Phloem ray
leaves             Pith                                     Xylem
                                                            ray



•Produced by                                                Primary
                                                            xylem

the vascular                                                      Secondary xylem
                                                                  Vascular cambium
                                                                   Secondary phloem
                                                                      Primary phloem

cambium and        Periderm
                                                                       First cork cambium       Cork




cork cambium       (mainly cork
                   cambia
                   and cork)

                   Primary
                   phloem
                   Secondary
                   phloem                                Secondary
                   Vascular                              xylem (two
                   cambium                               years of
                   Secondary                             production)
                   xylem                                        Vascular cambium
                                                                Secondary phloem                    Bark
                   Primary
                   xylem                                      Most recent
                   Pith                                       cork cambium                             Layers of
                                                                                      Cork             periderm
Secondary growth adds girth to stems
     and roots in woody plants
                Secondary phloem
                 Vascular cambium
             Late wood                                     Cork
Secondary                                                  cambium     Periderm
xylem        Early wood
                                                           Cork




                                                             Transverse section
                                                             of a three-year-
                                                             old Tilia (linden)
                               Xylem ray                     stem (LM)
                                                    Bark


    0.5 mm                                 0.5 mm
Secondary xylem

•As a tree             Growth ring
ages, the
older layers          Vascular
of                    ray

secondary
xylem no                Heartwood
               Secondary
longer         xylem     Sapwood
transport
water and               Vascular cambium

minerals                    Secondary phloem
                     Bark
                            Layers of periderm

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Plant Growth

  • 2. Meristems generate cells for new organs Primary growth in stems Shoot apical meristems Epidermis (in buds) Cortex Primary phloem Primary xylem Vascular cambium Lateral Pith Cork meristems cambium Secondary growth in stems Periderm Cork Pith cambium Cortex Primary Primary xylem phloem Root apical Secondary meristems xylem Secondary phloem Vascular cambium
  • 3. Woody plants Terminal bud Bud scale •In woody Axillary buds plants, primary Leaf scar This year’s growth Node and secondary (one year old) Stem growth occur Internode simultaneously One-year-old side branch formed but in different from axillary bud near shoot apex locations Last year’s growth Leaf scar (two years old) Scars left by terminal bud scales of previous winters Growth of two Leaf scar years ago (three years old)
  • 4. Primary Growth of Roots Cortex Vascular cylinder Epidermis Key Zone of Root hair maturation Dermal Ground Vascular Zone of elongation Apical Zone of cell meristem division Root cap 100 µm
  • 5. Primary Growth of Shoots Apical meristem Leaf primordia •A shoot apical meristem gives rise to a repetition of Developing vascular internodes strand and leaf- bearing nodes Axillary bud meristems 0.25 mm
  • 6. How a plant grows from a seed – primary growth • fuwopabecu.comoj.com
  • 7. Secondary growth adds girth to stems and roots in woody plants •Occurs in stems Primary and secondary growth in a two-year-old stem and roots of Epidermis woody plants Cortex Primary phloem Pith Primary xylem Vascular cambium but rarely in Vascular cambium Primary Primary phloem Cortex Epidermis xylem Phloem ray leaves Pith Xylem ray •Produced by Primary xylem the vascular Secondary xylem Vascular cambium Secondary phloem Primary phloem cambium and Periderm First cork cambium Cork cork cambium (mainly cork cambia and cork) Primary phloem Secondary phloem Secondary Vascular xylem (two cambium years of Secondary production) xylem Vascular cambium Secondary phloem Bark Primary xylem Most recent Pith cork cambium Layers of Cork periderm
  • 8. Secondary growth adds girth to stems and roots in woody plants Secondary phloem Vascular cambium Late wood Cork Secondary cambium Periderm xylem Early wood Cork Transverse section of a three-year- old Tilia (linden) Xylem ray stem (LM) Bark 0.5 mm 0.5 mm
  • 9. Secondary xylem •As a tree Growth ring ages, the older layers Vascular of ray secondary xylem no Heartwood Secondary longer xylem Sapwood transport water and Vascular cambium minerals Secondary phloem Bark Layers of periderm