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Photosynthesis: How Plants Harvest Light Energy
1.
© 2010 Pearson
Education, Inc. THE BASICS OF PHOTOSYNTHESIS • Photosynthesis: – Is used by plants, some protists, and some bacteria – Transforms light energy into chemical energy – Uses carbon dioxide and water as starting materials • The chemical energy produced via photosynthesis is stored in the bonds of sugar molecules.
2.
© 2010 Pearson
Education, Inc. Chloroplasts: Sites of Photosynthesis • Chloroplasts are: – The site of photosynthesis – Found mostly in the interior cells of leaves
3.
© 2010 Pearson
Education, Inc. • Inside chloroplasts are membranous sacs called thylakoids, which are suspended in a thick fluid, called stroma. • Thylakoids are concentrated in stacks called grana. • The green color of chloroplasts is from chlorophyll, a light- absorbing pigment.
4.
© 2010 Pearson
Education, Inc. • Stomata are tiny pores in leaves where carbon dioxide enters and oxygen exits.
5.
Leaf cross section Stomata Vein CO2
O2 Figure 7.2-1
6.
Leaf cross section Stomata Vein CO2
O2 Interior cell LM StromaGranum Thylakoid Chloroplast Outer membrane Inner membrane TEM Figure 7.2-2
7.
Leaf cross section Stomata Vein CO2
O2 Figure 7.2a
8.
StromaGranum Thylakoid Chloroplast Outer membrane Inner membrane TEM Figure
7.2b
9.
© 2010 Pearson
Education, Inc. The Overall Equation for Photosynthesis • In the overall equation for photosynthesis, notice that: – The reactants of photosynthesis are the waste products of cellular respiration.
10.
Carbon dioxide 6 O26 CO2 6
H2O C6H12O6 Water Glucose Photo- synthesis Oxygen gas Light energy Figure 7.UN1
11.
© 2010 Pearson
Education, Inc. • In photosynthesis: – Sunlight provides the energy – Electrons are boosted “uphill” and added to carbon dioxide – Sugar is produced
12.
© 2010 Pearson
Education, Inc. • During photosynthesis, water is split into: – Hydrogen – Oxygen • Hydrogen is transferred along with electrons and added to carbon dioxide to produce sugar. • Oxygen escapes through stomata into the atmosphere.
13.
© 2010 Pearson
Education, Inc. A Photosynthesis Road Map • Photosynthesis occurs in two stages: – The light reactions convert solar energy to chemical energy – The Calvin cycle uses the products of the light reactions to make sugar from carbon dioxide Blast Animation: Photosynthesis: Light-Independent Reaction
14.
Light H2O O2 Chloroplast Light reactions NADPH ATP Figure 7.3-1
15.
Light H2O O2 Chloroplast Light reactions NADPH ATP Calvin cycle CO2 NADP+ ADP P Sugar (C6H12O6) Figure 7.3-2
16.
© 2010 Pearson
Education, Inc. How Photosystems Harvest Light Energy • Light behaves as photons, discrete packets of energy.
17.
© 2010 Pearson
Education, Inc. • Chlorophyll molecules absorb photons. – Electrons in the pigment gain energy. – As the electrons fall back to their ground state, energy is released as heat or light.
18.
Light Photon Heat Light (fluorescence) Ground state Chlorophyll molecule Excited
state e– (a) Absorption of a photon Figure 7.8a
19.
© 2010 Pearson
Education, Inc. • A photosystem is a group of chlorophyll and other molecules that function as a light-gathering antenna.
20.
Chloroplast Figure 7.9-1
21.
Chloroplast Thylakoid membrane Pigment molecules Figure 7.9-2
22.
Chloroplast Thylakoid membrane Pigment molecules Antenna pigment molecules Primary electron acceptor Reaction- center chlorophyll
a Photon Electron transfer Reaction center Photosystem Transfer of energy Figure 7.9-3
23.
© 2010 Pearson
Education, Inc. How the Light Reactions Generate ATP and NADPH • Two types of photosystems cooperate in the light reactions: – The water-splitting photosystem – The NADPH-producing photosystem
24.
Primary electron acceptor Water-splitting photosystem Light H2O 2 H Reaction- center chlorophyll 2e– 2e– O2++ 1 2 Figure
7.10-1
25.
Primary electron acceptor Water-splitting photosystem Light H2O 2 H Reaction- center chlorophyll 2e– 2e– O2++ Energy to make ATP 1 2 Figure
7.10-2
26.
Primary electron acceptor Water-splitting photosystem Light H2O 2 H Reaction- center chlorophyll 2e– 2e– O2++ Energy to make ATP Primary electron acceptor 2e– Light NADPH-producing photosystem Reaction- center chlorophyll 2e– NADPH NADP 1 2 Figure
7.10-3
27.
© 2010 Pearson
Education, Inc. • The light reactions are located in the thylakoid membrane.
28.
© 2010 Pearson
Education, Inc. • An electron transport chain: – Connects the two photosystems – Releases energy that the chloroplast uses to make ATP
29.
Light H2O Thylakoid membrane 2e– O2 ATP NADP Light Stroma Inside thylakoid Photosystem Photosystem Electron
transport chain NADPH ADP P H+ ATP synthase To Calvin cycle H Electron flow H+ H H H H 1 2 Figure 7.11
30.
Light H2O Thylakoid membrane 2e– O2 ATP NADP Light Stroma Inside thylakoid Photosystem Photosystem Electron
transport chain NADPH ADP P ATP synthase To Calvin cycle Electron flow H 1 2 H H H H H H Figure 7.11a
31.
Water-splitting photosystem e– ATP NADPH e– e– e– e– e– e– NADPH-producing photosystem Figure 7.12
32.
THE CALVIN CYCLE:
MAKING SUGAR FROM CARBON DIOXIDE • The Calvin cycle: – Functions like a sugar factory within the stroma of a chloroplast – Regenerates the starting material with each turn © 2010 Pearson Education, Inc.
33.
Calvin cycle P P P Three-carbon moleculeRuBP sugar CO2
(from air) Figure 7.13-1
34.
ATP NADPH Calvin cycle ADP P NADP P P P P G3P
sugar Three-carbon moleculeRuBP sugar CO2 (from air) Figure 7.13-2
35.
ATP NADPH ADP P NADP P P P P P P G3P
sugar Three-carbon molecule G3P sugar G3P sugar RuBP sugar CO2 (from air) Calvin cycle Glucose (and other organic compounds) Figure 7.13-3
36.
ATP NADPH Calvin cycle ADP P NADP P P P P P P ADP
P ATP G3P sugar Three-carbon molecule G3P sugar G3P sugar RuBP sugar CO2 (from air) Glucose (and other organic compounds) Figure 7.13-4
37.
Evolution Connection: Solar-Driven Evolution •
C3 plants: – Use CO2 directly from the air – Are very common and widely distributed © 2010 Pearson Education, Inc.
38.
© 2010 Pearson
Education, Inc. • C4 plants: – Close their stomata to save water during hot and dry weather – Can still carry out photosynthesis • CAM plants: – Are adapted to very dry climates – Open their stomata only at night to conserve water
39.
Sugar C4 Pathway (example: sugarcane) C4
plant Calvin cycle Cell type 1 Four-carbon compound Cell type 2 CO2 CO2 Figure 7.14a
40.
CAM plant Sugar Calvin cycle Day Four-carbon compound Night CAM Pathway (example:
pineapple) CO2 CO2 Figure 7.14b
41.
Carbon dioxide 6 O26 CO2 6
H2O C6H12O6 Water Glucose Photo- synthesis Oxygen gas Light energy Figure 7.UN1
42.
Light reactions CO2 O2 H2O (C6H12O6) NADPH Light Sugar ATP ADP P NADP Calvin cycle Figure 7.UN2
43.
Light reactions CO2 O2 H2O (C6H12O6) NADPH Light Sugar ATP ADP P NADP Calvin cycle Figure 7.UN3
44.
Carbon dioxide 6 O26 CO2 6
H2O C6H12O6 Water Glucose Photosynthesis Oxygen gas Light energy Figure 7.UN4
45.
Light H2O O2 Chloroplast Light reactions NADPH ATP Calvin cycle CO2 NADP+ ADP P Sugar (C6H12O6) Stack of thylakoids
Stroma Figure 7.UN5
46.
acceptor Water-splitting photosystem Photon H2O 2 H Chlorophyll 2e– O2++ ATP NADPH-producing photosystem Chlorophyll NADPH NADP+ e– 2e– 2e– 2e– e– acceptorADP Photon 2 1 Figure 7.UN6
47.
NADPH Calvin cycle ADP P NADP P ATP G3P CO2 Glucose and other
compounds Figure 7.UN7
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