3. 1. IntroductIon
An inflorescence is
a group or cluster
of flowers arranged on a stem that is
composed of a main branch or a
complicated arrangement of branches.
Strictly, it is the part of the shoot of seed
plants where flowers are formed and
which is accordingly modified. The
modifications can involve the length and
the nature of the internodes and
the phyllotaxis(arrangement of leaves), as
well as variations in the proportions,
compressions,swellings, adnations (union
of unlike parts), connations (fusion of
similar organs) and reduction of main and
6. The
stem holding the whole
inflorescence is called
a peduncle and the main stem
holding the flowers or more
branches within the inflorescence
is called the rachis. The stalk of
each single flower is called
a pedicel.
The fruiting stage of an
inflorescence is known as
an infructescence.
7.
8. 2. General characterIstIcs
Inflorescences
are described by many different
characteristics including how the flowers are
arranged on the peduncle, the blooming order of
the flowers and how different clusters of flowers
are grouped within it. These terms are general
representations as plants in nature can have a
combination of types.
9. 2.1. Bracts
Inflorescences
usually have modified
foliage different from
the vegetative part of the plant.
Considering the broadest meaning of
the term, any leaf associated with an
inflorescence is called a bract. A bract
is usually located at the node where
the main stem of the inflorescence
forms, joined to the main stem of the
plant, but other bracts can exist within
10. According to the presence or absence
of bracts and their characteristics can
be distinguished in:
2. Ebracteate inflorescences: No
bracts in the inflorescence.
11. 2. Bracteate inflorescences: The
bracts in the inflorescence are very
specialized, sometimes reduced to
small scales, divided or dissected.
12. 3. Leafy inflorescences: Though often reduced in
size, the bracts are unspecialized and look like the
typical leaves of the plant, so that the
term flowering stem is usually applied instead of
inflorescence. This use is not technically correct,
as, despite their 'normal' appearance,
these leaves are considered, in fact, bracts, so
that 'leafy inflorescence' is preferable.
14. 2.2. Terminal flower
Plant
organs can grow according to two
different schemes,
namely monopodial or racemose( growth
habits grow upward from a single
point) and sympodial or cymose( growth
have a specialized lateral growth pattern
in which the apical meristem is
terminated). In inflorescences these two
different growth patterns are called
indeterminate or determinate, and
15.
16. Determinate
inflorescence: Sympodial growth. The
terminal bud forms a terminal flower and
then dies out. Other flowers then grow
from lateral buds. In determinate
inflorescences the terminal flower is
usually the first to mature (precursive
development), while the others tend to
mature starting from the bottom of the
stem. This pattern is
called acropetal maturation. When
flowers start to mature from the top of
18. Indeterminate
inflorescence: Monopodial growth. The
terminal bud keeps growing and
forming lateral flowers. A terminal
flower is never formed.
In indeterminate inflorescence there is
no true terminal flower and the stem
usually has a rudimentary end. In
many cases the last true flower formed
by the terminal bud
(subterminal flower) straightens up,
appearing to be a terminal flower.
21. 1.4. meTaToPy
Metatopy
is the placement of organs out of
their normally expected position: typically
metatopy occurs in inflorescences when
unequal growth rates alter different areas of
the axis and the organs attached to the axis.
When a single or a cluster of flower(s) is
located at the axil of a bract, the location of
the bract in relation to the stem holding the
flower(s) is indicated by the use of different
terms and may be a useful diagnostic
22. Typical placement of bracts include:
Some plants have bracts that subtend the
inflorescence, where the flowers are on
branched stalks; the bracts are not
connected to the stalks holding the flowers,
but are adnate or attached to the main stem
(Adnate describes the fusing together of
different unrelated parts. When the parts
fused together are the same, they are
connately joined.)
Other plants have the bracts subtend
the pedicel or peduncle of single flowers.
23. Metatopic
placement of bracts include:
When the bract is attached to the stem
holding the flower (the pedicel or
peduncle), it is said to be recaulescent;
sometimes these bracts or bracteoles
are highly modified and appear to be
appendages of the flower calyx.
Recaulescences is the fusion of the
subtending leaf with the stem holding the
bud or the bud itself, thus the leaf or
bract is adnate to the stem of flower.
When the formation of the bud is shifted
up the stem distinctly above the
25. 3. OrganizatiOn
The
main groups of
inflorescences are
distinguished by branching.
Within these groups, the most
important characteristics are
the intersection of the axes
and different variations of the
model.
26. 3.1 Simple
inflOreScenceS
3.1.1 indeterminate
Indeterminate simple inflorescences
are generally called racemose. The
main kind of racemose
inflorescence is the raceme. The
other kind of racemose
inflorescences can all be derived
from this one by dilation,
compression, swelling or reduction
28. A spike is
a type of raceme with flowers that
do not have a pedicel.
Spike
Plantago media
(spike)
29. A
racemose corymb is an unbranched,
indeterminate inflorescence that is flat-topped
or convex due to their outer pedicels which are
progressively longer than inner ones.
Racemose corymb
Iberis umbellata
(racemose
30. An umbel is a type of raceme with a short axis
and multiple floral pedicels of equal length that
appear to arise from a common point.
Umbel
Astrantia minor
(umbel)
31. A spadix is a spike of flowers densely arranged
around it, enclosed or accompanied by a highly
specialised bract called a spathe. It is
characteristic of the Araceae family.
Spad
ix
Arum maculatum
(spadix)
32. A flower head or capitulum is a very
contracted raceme in which the single sessile
flowers share are borne on an enlarged stem
Head
(round)
Dipsacus
fullonum(head)
33. A catkin or ament is a scaly, generally drooping
spike or raceme. Cymose or other complex
inflorescences that are superficially similar are
also generally called thus.
Catkin (racemose
or spicate)
Alnus incana (ament)
35. Cymes are further divided
according to this scheme:
Secondary buds always develop on the same
side of the stem: helicoid cyme or bostryx.
Bostryx
(lateral and
Hypericum
perforatum (bostryx)
36. The successive pedicels are aligned
on the same plane: drepanium
Drepanium (lateral Gladiolus
and top view)
imbricatus
37. The successive pedicels are
arranged in a sort of
spiral: cincinnus
Cincinnus (lateral Symphytum
officinale
and top view)
38. The successive pedicels follow a zig-zag
path on the same plane: rhipidium
Rhipidium
(lateral and top Canna sp. (rhipidium
view)
39. A cyme can also be so compressed that it looks
like an umbel. Strictly speaking this kind of
inflorescence could be called umbelliform
cyme, although it is normally called simply
'umbel'.
Pelargonium
Umbelliform cymezonale
40. Another kind of definite simple inflorescence is
the raceme-like cyme or botryoid; that is as a
raceme with a terminal flower and is usually
improperly called 'raceme'.
Botryoi
d
Berberis
vernae (botryoid)